**580**wrote:

Hi,

I am trying to find a formal reference and more details to this equation:

I saw it in this Nature Genetics paper (Online Methods section), by Park *et al.* (2010).
This equation determines the contribution of a SNP to the genetic variance of a trait or effect size (ES), taking into account its regression effect (Beta) and allele frequencies (f).

I have searched in books, articles and throughout the internet, but found nothing.

Could someone provide a more detailed reference or explanation about it?

Thank you!