I downloaded a file from enhancer database (Slidebase) named "enhancer tss associations". However, I am facing a problem in identifying what does coordinates in the first three columns of this bed file represents. I am aware that the fourth column containing entries like "chr1:167440766-167441089; NM_052862; RCSD1; R:0.319; FDR:0" means enhancer coordinates,Transcript accession number,gene symbol,some_score, and False Discovery rate. I am not sure what kind of score does R score represents. I went through the paper of andersson et al to which the website points. However, I couldn't find anything. Also, the last two columns don't make sense to me.